Prof. Eva Jablonka

מכון להסטוריה ופילוסופיה סגל אקדמי בכיר
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Phone: 03-6408910
Another phone: 03-6409198
Fax: 03-6409463
Office: Gilman-humanities, 399


Born:  Poland, 9th January 1952. Immigrated to Israel 1957.



Research Interests

Heredity, Epigenetics. Evolutionary Biology, Behavioural Ecology, Developmental Biology, Language and Cultural evolution, History of Genetics and the Philosophy of Science, Evolution of mind and of nervous systems 


EMBO (European Molecular Biology Organisation) Post-doctoral Fellowship (10.87 – 01.88).

Post-doctoral Fellowship in the Edelstein Centre for the History and Philosophy of Science, Technology and Medicine,HebrewUniversity,Jerusalem(01.88 – 01.89).

Allon Fellowship,Tel-AvivUniversity(10.90 – 10.93).

Fellow of the CollegiumBudapest, Institute for Advanced Studies (10.94 – 2.95).

Fellow of the Institute for Advanced Studies,Berlin  (10.97 – 7.98).

Visiting Fellow in All Souls College, Oxford, Trinity Semester 2011.

Academic Qualifications

1976    B.Sc. in Biology fromBen-Gurion University,Israel.

1980    M.Sc. (with distinction) in Microbiology fromBen-Gurion University,Israel, for work carried out 1978-80.  Subject of Thesis: Regulation of Peptide Transport in E. coli.

1988    Ph.D. in Genetics from the Hebrew University, Jerusalem, Israel, for work carried out 1983-87 under the supervision of Prof. Menashe Marcus and Prof. Howard Cedar.  Subject of Thesis: Alterations in Chromosomal Structure and Genic Activity in the Inactive X chromosome in Female Mammals.


Teaching and Research Positions

1978 - 1980     Assistant in Genetics, Microbiology and Biochemistry courses inBenGurionUniversity.

1981 - 1982     Research work for the Van-Leer Institute,Jerusalem. Subject: The Biological Perspective in the Work of Jean Piaget.

1987    Post-doctoral position in the Medical Research Council Mammalian Development Unit, London, with Dr. Anne Mclaren and Dr. Marilyn Monk.

1988 - 1990     Post-doctoral fellowship in the Edelstein Centre for the History and Philosophy of Science Technology and Medicine.

1987 - 1989     Taught the following: (i) A course in ‘Genetic Analysis’ for the students of the “Interdisciplinary Project for Promoting Excellence” in Tel-Aviv University; (ii) An M.Sc. course ‘Philosophical Problems in Biology’, in Tel-Aviv University; (iii) An M.Sc. course in the Philosophy of Biology at the Hebrew University; (iv) A course in the History of Genetics for M.Sc. students in Tel-Aviv University.

1986 - 1989     Together with Lia Ettinger created and organized the Discussion Group in Theoretical Biology at theHebrewUniversity.

1990 - 1991     Fellow in the Interdisciplinary Group on ‘Biological Foundations of Human Culture’ in Bielefeld University,Germany.

1990 - 1993     Lecturer, The Cohn institute for History and Philosophy of Science and Ideas, teaching Genetics, Evolutionary Biology (in the Zoology Department), Philosophy of Biology, History of Genetics.

1993 – 2000    Tenured senior lecturer,Tel-Aviv University.

1994 - 1995     Fellow of the Collegium Budapest Institute of Advanced Studies. Member of an International Theoretical Biology Group.

1997 - 1998     Fellow of the Institute for Advance Studies inBerlin.

2000-2005   Associate Professor,Tel-AvivUniversity

2000-2001    Visiting Scholar (Sabbatical) in The Museum of Vertebrate Zoology, University of California, Berkeley.

2005- present Full professor, Tel-Aviv University.

2011 (Trinity term) Visiting Fellow, All Souls College, Oxford.

2011 (September-October) Visiting professor in the University of Perpignan, France, department of  Ecology and Evolution (with Professor Christoph Grunau).

2013 Course on “The Extended Evolutionary Synthesis: An Epigenetic Approach” in Fiucruz public health department, Rio de Janeiro


Books in English

Eva Jablonka and Marion J. Lamb (1995) Epigenetic Inheritance and Evolution – the Lamarckian Dimension,OxfordUniversity Press. (Second printing with new introduction 1999).

Eytan Avital and Eva Jablonka. (2000) Animal Traditions: Behavioural Inheritance in Evolution.CambridgeUniversity Press. (Paperback edition 2005)

Eva Jablonka and Marion L. Lamb (2005) Evolution in Four Dimensions. MIT Press. (Paperback edition 2006, Italian translation 2007, Hebrew translation 2008, Slovenian 2009 Portuguese 2010, Turkish 2011; Spanish 2013); second revised edition due 2014.

Snait Gissisand Eva Jablonka (Editors) (2011) The Transformations of Lamarckism: From Subtle Fluids to Molecular Biology. MIT Press

Book in Hebrew:

Eva Jablonka (1994) (In Hebrew) History of Heredity.  Ministry of Defence Publishing House, Israel.

Eva Jablonka (1994 -1997) (In Hebrew) Evolution—A Textbook in Evolutionary Biology for the Open University, Israel. Open University Press. 7 units. 700 pages.


Papers and chapters in books (English)

Tavori H*., Kimmel Y., and Barak Z. (1981) Toxicity of leucine-containing peptides in Escherichia coli caused by circumvention of leucine transport regulation.  Journal of  Bacteriology,146, 676-683. (*Jablonka E.)

Barak Z., Gollop N., and Tavori H.* (1981) Peptide uptake - a method circumventing regulation of leucine transport in E. coli.  13th FEBS Meeting, 1981. (*Jablonka E.)

Barak Z., Tavori H.*, and Gollop N. (1981) Negative control of leucine transport in E. coli  K-12.  Israel Journal of Medicine. (*Jablonka E.)

Gollop N., Tavori H.*, and Barak Z. (1982) Acetohydroxy acid synthase is a target for leucine-containing peptide toxicity in Escherichia coli.  Journal of Bacteriology, 149, 387-390. (*Jablonka E.)

Jablonka-Tavory E.* (1982) Genocopies and the evolution of interdependence.  Evolutionary Theory, 6, 167-170. (*Jablonka E.)

Jablonka E., Goitein R., Marcus M., and Cedar H. (1985) DNA hypomethylation causes an increase in DNase-I sensitivity and an advance in the time of replication of the entire inactive X chromosome.  Chromosoma, 93, 152-156.

Jablonka E., Goitein R., Sperling K., Cedar H., and Marcus M. (1987) 5-aza-C-induced changes in the time of replication of the X chromosomes of Microtus agrestis are followed by non-randon reversion to a late pattern of replication.  Chromosoma, 95, 81-88.

Jablonka E., Goitein R., Sperling K., Cedar H., and Marcus M. (1987) Regulatory sites determining the time of replication of the X chromosome of Microtus agrestis7th International Congress of Human Genetics, Part I.

Jablonka E., and Lamb M.J. (1988) Meiotic pairing constraints and the activity of sex chromosomes.  Journal of Theoretical Biology, 133, 23-36.

Jablonka E., and Lamb M.J. (1989) The inheritance of acquired epigenetic variations.  Journal of Theoretical Biology, 139, 69-83.

Ettinger L., Jablonka E., and MacLaughlin P. (1990) On the adaptations of organisms and the fitness of types.  Journal of Philosophy of Science, 57, 499-513.

Jablonka E., and Lamb M.J. (1990) The evolution of heteromorphic sex chromosomes. Biological Reviews, 65, 249-276.

Jablonka E., and Lamb M.J. (1990) Lamarckism and Aging. Gerontology, 36, 323-332.

Ettinger L., R. Falk, and Jablonka E. (1991) On causality, heritability and fitness. Biology and Philosophy, 6, 27-29.

Jablonka E., and Lamb M.J. (1991) Sex chromosomes and speciation. Proceeding of the Royal Society London B., 243, 203-208.

Jablonka E. and Lamb M. J. (1992) Species and Speciation. Nature, 356, 752.

Jablonka E., Lachmann M. and Lamb M. J. (1992) Evidence, mechanisms and models of the inheritance of acquired characters. Journal of Theoretical Biology. 158, 245-268.

Avital E. and Jablonka E. (1994) Social learning and the evolution of behaviour.  Animal Behaviour, 48, 1195-1199.

Jablonka E. (1994) Inheritance systems and the evolution of new levels of individuality. Journal Theoretical Biology, 170, 301-309.

Jablonka E. and Szathmáry E. (1995). The evolution of information storage and heredity.  Trends in Ecology and Evolution, 10, 206-211. 

Jablonka E. and Regev A. (1995) Gene number, methylation and biological complexity. Trends in Genetics, 11, 383-4.

Jablonka E., Oborny B, Molnár E., Kisdi E., Hofbauer J., and Czárán T. (1995) The adaptive advantage of phenotypic memory. Philosophical Transactions of the Royal Society, London B., 350, 133-141.

Avital E. and Jablonka E. (1996) Adoption, memes and the Oedipus complex: a reply to Hansen. Animal Behaviour, 51, 476-477.

Lachman M. and Jablonka E., (1996) The inheritance of phenotypes: an adaptation to fluctuating environment. Journal of Theoretical Biology, 181, 1-9

Jablonka E. and Rechav G. (1996) The evolution of language in the light of the evolution of literacy. In: The Major Origins of Language, (Ed. J. Trabant). Collegium Budapest, pp. 70-88.

Jablonka E. (1996) Do cells show off?  Somatic selection and the nature of inter-cellular signalling. Trends in Ecology and Evolution, 11, 395-396.

Falk R. and Jablonka E. (1997) Inheritance: transmission and development. In: Human by Nature: Between Biology and the Social Sciences (Eds: Weingart P., Mitchell S.D., Richerson P.J. and Maasen S.) Lawrence Erlbaum Associates: Mahwah, NJ. pp. 390-400.

Jablonka E. (1998) The Evolution of Inheritance and Environment. In: The Co-Action between living Systems and the Planet.  (Eds.: Greppin, H., Agosti, R.D., and Penel C. ), pp. 131-145.  University of Geneva.

Jablonka E. and Lamb M.J. (1998) Epigenetic inheritance in evolution (A target article). Journal of Evolutionary Biology, 11, 159-183.

Jablonka E. and Lamb M.J. (1998) Genic-Neo Darwinism—is it the whole story? Journal of Evolutionary Biology, 11,  243-260.

Jablonka E. and Lamb M.J. (1998) Bridges between evolution and development. Biology and Philosophy, 13, 119-124.

Avital E., Jablonka E., and Lachmann M. (1998) Adopting adoption. Animal Behaviour, 55: 1451-1459.

Regev A., Lamb M.J., and Jablonka E. (1998) The role of DNA methylation in invertebrates: developmental regulation or genome defense?  Molecular Biology and Evolution, 15, 880-891.

Jablonka E., Lamb M.J., and Avital E. (1998). Lamarckian mechanisms in Darwinian evolution. Trends in Ecology and Evolution, 13, 206-210.

Lachmann M., Sella G., and Jablonka E.  (2000) On the advantages of Information sharing. Proc.Roy.Soc.  B 267, 1287-1293.

Jablonka E. and Ziman J.  (2000)  Biological evolution: processes and phenomena.  In: Technological Innovation as an Evolutionary Process. (Ed: Ziman J.).pp. 13-27. Cambridge University Press, Cambridge.

Jablonka E.   (2000) Lamarckian inheritance systems in biology: a source of metaphors and models in technological evolution. In: Technological Innovation as an Evolutionary Process. (Ed: Ziman J.), pp.27-41. Cambridge University Press, Cambridge.

Dor D. and Jablonka E.  (2001) From cultural selection to genetic selection: a framework for the evolution of language. Selection, 1, pp. 33-57.

Jablonka E.  (2001) The systems of inheritance.  In: Cycles of Contingency (Eds: Oyama, S., Griffiths P. and Gray R., MIT Press), pp 99-116.

Daniel Dor and Jablonka Eva (2001). How language changed the genes. In  Trabant  J.  Ward. S. (editors). New Essays on the Origin of Language. Mouton de Gruyer: Berlin, pp 149-175.

Eva Jablonka and Marion J. Lamb  (2002) “Epigenetic inheritance”, an entry for the Encyclopaedia of Social Sciences. Vol. 7  pp 4706-4710.

Jablonka E. and Lamb M.J. (2002) Epigenetics. In: Encyclopedia of Evolution, Vol. 1 (Ed. Pagel, M.), pp. 310-311. Oxford University Press, Oxford.

Jablonka E. and Lamb M.J. (2002) Lamarckism. In: Encyclopedia of Evolution, Vol. 2 (Ed. Pagel, M.), pp. 602-605. Oxford University Press, Oxford.

Jablonka E. and Lamb M.J. (2002) Creating bridges or rifts?  Developmental systems theory and evolutionary developmental biology. BioEssays, 24, 290-291.

Jablonka E., Matzke, M., Thieffry, D. and Van Speybroeck, L. (2002) The genome in context: biologists and philosophers on epigenetics. BioEssays, 24, 392-394.

Jablonka E. 2002.  Information: its interpretation, its inheritance and its sharing. Philosophy of Science, 69: 578-605.

Jablonka E. (2002) Between Development and evolution: how to model cultural change. Proceedings of the British Academy, 112: 27-41.

Jablonka E. and Lamb M.J. (2002) The changing concept of epigenetics. Annals of the New York Academy of Sciences, 981: 82-96

Jablonka Eva  (2004) From Replicators to Heritably Varying Phenotypic Traits: The Extended Phenotype Revisited. Biology and Philosophy, 19: 353-75.

Jablonka Eva (2004) Epigenetic Epidemiology. International Journal of Epidemiology 33:929–35.

Jablonka Eva (2004). The peculiarities of mammalian sex chromosomes: an epigenetic view. BioEssays, 26(12): 127-32.

Dor Daniel and Jablonka Eva (2004)  Culture and Genes in the Evolution of Human Language In: Goren-Inbar, Naama, and John D. Speth (eds). Human Paleoecology in the Levantine Corridor. Oxford, England: Oxbow Press, pp.105-115.

Jablonka Eva and Marion Lamb(2006) The evolution of information in the major transitions. Journal of Theoretical Biology 239: 236-246.

Jablonka Eva and Marion Lamb( 2006) Evolutionary epigenetics,  In Evolutionary Genetics: Concepts and Case Studies, (eds CW Fox and JB Wolf), Oxford University Press, New York, pp 252-264.

Jablonka Eva (2006) Commentary: Induction and selection of variations during cancer development. International Journal of Epidemiology doi:10.1093/ije/dyl188

 Jablonka Eva (2006) Le darwinisme évolue aussi LA RECHERCHE, Nº 396 46 - 50 (in French)

Jablonka Eva  (2006) Genes as followers in evolution – a post-synthesis synthesis? Biology and Philosophy, 21: 143-154.

Jablonka Eva (2007) Epigenetics and Evolution: An Overview Accademia Nazionale delle Scienze detta dei XL Memorie di Scienze Fisiche e Naturali 124° (2006), Vol. XXX, t. I, pp. 1-11

Eva Jablonka  (2007) Information is everything that can be interpreted. In: Interact or Die. V2_ Publishing/NAi Publishers 2007, pp 28-43. (Interview)

Eva Jablonka (2007) The developmental construction of heredity. Developmental psychobiology 49: 808-817.

Eva Jablonka and Marion Lamb(2007) The expanded evolutionary synthesis – a response to Godfrey-Smith, Haig, and West-Eberhard.  Biology and Philosophy Biol. Philos. 22: 453-472.

Eva Jablonka and Marion Lamb(2007) Précis of: Evolution in Four Dimensions.  Brain and Behavioral Science 30: 353-365.

Eva Jablonka and Marion Lamb(2007) Bridging the Gap: The Developmental Aspects of Evolution.  Brain and Behavioral Science 30: 378-392.

Eva Jablonka and Marion Lamb(2007) Reply to Wilkins on review of Evolution in Four Dimensions BioEssays 29:308–309

Simona Ginsburgand Eva Jablonka (2007) The Transition to Experiencing: I. Limited Learning and Limited Experiencing. Biological Theory. 2(3) 218–230.

Simona Ginsburgand Eva Jablonka (2007) The Transition to Experiencing: II. The Evolution of Associative Learning Based on Feelings. Biological Theory 2(3) 231–243.

Eva Jablonka (2008) The Five Mothers: Heredity and Evolution from a Developmental Perspective.  In the Proceedings of Baxter lectures: The Futures of Darwinism (In Italian).

Ehud Lamm and Eva Jablonka (2008) The nurture of nature: hereditary plasticity in evolution. Philosophical Psychology 21:305-319.

Ehud Lamm and Eva jablonka (2008) Review Essay: Integrating Evolution and Development.  Perspectives in Biology and Medicine 51:636-647.

Eva Jablonka and Marion Lamb (2008) The epigenome in evolution: beyond the modern synthesis. Вестник ВОГиС 12: 242-254.

Eva Jablonka (2008) Soft Inheritance: Challenging the Modern Synthesis Genetics and Molecular Biology 31:389-395.

Eva Jablonka and Marion Lamb (2008) The Return of Soft Inheritance. Seed  December 2008 pp 25-26

Eva Jablonka and  Gal Raz (2009) Transgenerational Epigenetic Inheritance: Prevalence, Mechanisms, and Implications for the Study of Heredity and Evolution. Quarterly Review of Biology 84: 131-176

Eva Jablonka (2009) In: Evolutionary Theory: 5 Questions (Eds: Oftedal G., Berg O. Friis JK, Rossel P. abd Norup MS).  Automatic Press. Pp.63-77 

Eva Jablonka and Marion Lamb (2009) The Pillars of Darwinism.  Project Syndicate (February 2009

Simona Ginsburg and Eva Jablonka (2009) Epigenetic Learning in Non-Neural Organisms. Journal of Bioscience 34(4):633-46.

Eva Jablonka and Marion Lamb (2010) Transgenerational epigenetic Inheritance. In: Evolution, The Extended Synthesis. (eds Pigliucii M and Müller GB).  pp 137-174. MIT Press.

Simona Ginsburg and Eva Jablonka (2010) Experiencing: a Jamesian approach   Journal of Consciousness Studies 17:102-124.

Simona Ginsburg and Eva Jablonka (2010) Associative learning: a factor in the Cambrian explosion. Journal of Theoretical Biology 266:11–20.

 Daniel Dor and Eva Jablonka (2010) Canalization and plasticity in the evolution of linguistic communication In The Evolution of Human Language (Eds Larson RK, Deprez V. and Yamakido H.), pp135-147 Cambridge University Press.

Omri Tal, Eva Kisdi and Eva Jablonka (2010) Epigenetic contribution to covariance between relatives. Genetics 184:1037–1050.

Jablonka Eva and Marion J. Lamb (2011) Changing thought styles: the concept of soft inheritance in the 20th century.  Vérité, Widerstand, Development: At Work with / Arbeiten mit / Travailler avec Ludwik Fleck, hg. v. Rainer Egloff und Johannes Fehr, Zürich 2011 (Collegium Helveticum Heft 12). Pp. 119-157.

Eva Jablonka and Ehud Lamm (2011) The Epigenotype: a dynamic network-view of development. International Journal of Epidemiology International Journal of Epidemiology  41(1):10-13.

Jablonka E and Ginsburg Simona. (2012) Scaffolding Emotions and Evolving Language. Behavioral and Brain Sciences. 35(3):154-5.

Iddo Tavory, Simona Ginsburg and Eva Jablonka (2012) Culture and Epigenesis: A Waddingtonian view. The Oxford Handbook of Culture and Psychology (ed. Jaan Valsiner).Chapter 30,  pp. 662-676

Eva Jablonka, Simona Ginsburg and Daniel Dor (2012). The Co-evolution of language and emotions. Philosophical transactions of the Royal Society B, 367: 2152-2159. doi:10.1098/rstb.2012.0117

Eva Jablonka (2013) Behavioral epigenetics in ecological context.  Behavioral Ecology. doi: 10.1093/beheco/ars115

Eva Jablonka (2013) Epigenetic Plasticity: The Responsive Germline. Progress in Biophysics and Molecular Biology. 111, 99- 107.

Eva Jablonka (2013) Epigenetic Variations in Heredity and Evolution. Clinical Pharmacology and Therapeutics. doi:10.1038/clpt.2012.158

Eva Jablonka (2013) Some Problems with Genetic Horoscopes. In: Genetic Explanations: Sense and Nonsense. Pp 71-80. Harvard University Press.

Mesoudi, A., Blanchet, S., Charmantier, A., Danchin, E., Fogarty, L., Jablonka, E., Laland, K.N., Morgan T.J.H., Müller, G.B., Odling-Smee, F. J., Pujol, B. (2013) Is non-genetic inheritance just a proximate mechanism? A corroboration of the Extended Evolutionary Synthesis. Biological Theory 7(3)

Laor Yuval and Jablonka Eva (2013) The Evolution and Development of Culture. History and Theory,  52 (2) 290-299.

Simona Ginsburg and Eva Jablonka (in press) Memory, imagination and the evolution of modern language. In:  Social Origins of Language. Oxford University Press.

Iddo Tavory, Simona Ginsburg and Eva Jablonka (in Press) The reproduction of the social: a Developmental system view. Linnda Caporael, James Griesemer and William Wimsatt (eds) Scaffolding in Evolution, Culture and Cognition. MIT Press.

Eva Jablonka and Marion Lamb (in press) Disturbing Dogmas: Biologists and the History of Biology. Science in Context.

Daniel Dor and Eva Jablonka (in press) Why we need to move from gene-culture co-evolution to culturally-driven co-evolution. In: Social Origins of Language. Oxford University Press.

Bronfman Zohar, Simona Ginsburg and Eva Jablonka (in press) The epigenetics of neural learning. In: Handbook of Cognitive neuroscience of associative learning.

Simona Ginsburg and Eva Jablonka (in press) The Teleological Transitions in Evolution: a Gántian View. Systems Chemistry.

Reviewed papers and book chapters in Hebrew

Eva Jablonka (1994) The significance of epigenetic inheritance systems. Iyun, 42, 533-539. (in Hebrew).

Eva Jablonka (2007) Radical conservatism in Leibowitz’s Philosophy of biology. In Yehsayhu Leibowitz: Between Conservatism and Radicalism. Reflections on His Philosophy. Ed: Aviezer Ravitsky. The Van Leer Jerusalem Institute/Hakibbutz Hemeuchad Publishing House. Pp.190-204 (in Hebrew)

 Selected Book Reviews (since 2000)

1. Selection – The Evolution of the Common Good.  Selection 1-3.  pp 247-250. 2000.

2. Journal of Heredity  - Thinking about Evolution. Volume 87, pp.  125-6.  2001.

3. Acta Biothertica (with Arnon Levy) – Marcello Barbieiri (2003). The Organic codes: an introduction to semantic biology, 52(1) 65-9). 2004.

4. Biology and Philosophy – (with Eytan Avital) Animal innovation Biology and Philosophy, 21: 135-147,  2006

5. Consciousness Studies  -  (with Simona Ginsburg) A book review of Derek Denton's book, The Primordial Emotions: The Dawning of Consciousness . Consciousness Studies 19:  105-109,  2006

6. BioEssays  - The nervous system in development and evolution. BioEssays 31:687–689, 2009.

7. . Consciousness Studies -  (with Simona Ginsburg) A book review of Thomas Nagel’s book Mind and Cosmos: Why the Materialistic Neo-Darwinian Conception of Nature is Almost Certainly Wrong. Consciousness Studies, 20, No. 9–10, pp. 177–205, 2013. 


1981    Awarded the Landau Prize of Israel for outstanding M.Sc. work.

1988    Awarded Marcus prize for outstanding Ph.D work.

1990    Awarded the Allon Fellowship in 1990.


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